Penn State football has never beaten these FBS teams

The Penn State Nittany Lions have beaten 85 different FBS teams in their 130-year history, but these eight teams have beaten Penn State and the Nittany Lions have never gotten revenge.
Mississippi Rebels quarterback Jaxson Dart (2)
Mississippi Rebels quarterback Jaxson Dart (2) / Brett Davis-USA TODAY Sports
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The 2000 season was one of Penn State’s most disappointing years. It began with a 29-5 loss to USC at a neutral site to open the year, but then the Nittany Lions returned home to face Toledo for the first time, and things went from bad to worse. 

Toledo handled Penn State 24-6 and the teams have not played since. The Nittany Lions started the season 1-4 after beating Louisiana Tech and losing to Pitt and Ohio State. Joe Paterno’s team finished the year 5-7, the first of four losing seasons in five years to start the 21st century.